I know there is R9 280X with BIOS switch--did you accidentally flip the switch to the non-Mac-EFI ROM while prepping the MP for sale?
Did you change cables or monitors and start using a different port, perhaps one that hasn't been providing boot screens all along?
Also there's also the possibility if you short the correct pins that you can restore to factory flash (to recover from a bad flash). But that is difficult to do on purpose, much less accidentally, so that seems extraordinarily unlikely.
Personally I can't think of any other reason a flashed Mac EFI would spontaneously stop providing boot screens.