so what makes the S15 a $1,400 MSRP unit when these others (that aren't labeled "AV") go for $450?
It was a $1400 MSRP unit way back in the day. It certainly isn't worth anything near that now.
What made it so valuable were the extra features designed specifically for audiophiles (snobby home theater customers). These features are not necessary for Mac Pros, but do no harm for snobby Apple customers.