Depends.
Apple actually killed distributed compilation in the last version of XCode, so you can't send compiling to servers any more. If Apple kills the Mac Pro, developers can't even build Mini clusters to augment XCode any more.
On the other hand, Apple is allowed to cheat. No reason they can't have a bunch of generic PCs running OS X. That's exactly what they did while they were writing OS X for Intel.
Another possibility is that Apple makes it 'easy' for iOS developers through some sort of Cloud-ification ... this could be an "Included in the Package" sweetener that's part of Apple's "Developer ID Card" (which will probably carry an annual fee) bit that's going to be coming to the Mac.
BTW, thanks for the correction on the 500MHz G4; my recollection had been that some quantity of them had supposedly shipped before it was canned.
Except most tasks are a real pain to be done in "the cloud". I work on a Mac Pro and use a cluster for scientific research, and... ... if its at all like what I deal with, and what I've heard from other software engineers, the workstation is still a very critical part of the computing toolkit. And if Apple needs a workstation internally, they probably will just want to build them and sell them to us too.
That's kind of the direction I was thinking. My expectation is that Apple could work out these headaches for whatever they want to roll-out and have it end up being basically as user-friendly as the photo sharing bit currently is on iOS.
Of course, the big difference I see is that Apple's not going to be stuck with some slow & thin pipe to their own server farm. I already see this when watching Apple's bloated website load its pages...it probably appears to be just fine while they're working on it in-house on a fast machine & fast connection.
Apple is heavily invested in OpenCL (among other high-end technologies).
Anybody at Apple doing OpenCL needs what I need: a Mac Pro with a fat GPU.
I've been monitoring for news on Grand Central Dispatch and OpenCL...its important stuff to the higher end of the power scale...but that frontier has seemed to be pretty quiet, for far too long.
-hh