Origin is a download store for the Feral games, it has a few nice features like save game sync but it is purely a download service like the Feral Download Store or MacGameStore. It does not embed DRM into the game like Apple and Valve do with the Mac AppStore and Steam. The game is the same as you would get on a DVD or from the feral download store.
You can delete Origin if you wish and the game will keep working, you can even copy the game to your second Mac and activate it without installing Origin again.
The only server connection is to validate your product key and the game only does this when you first activate the game on new hardware, after that it does not phone home to check the DRM without your specific request to do so via an "update" or "deactivate" button.
You can have it installed on up to 5 Macs at once (you can add and remove Macs) and activating multiple times on the same Mac only counts as a single activation.
Edwin
Origin is a download store for EA games. Origin recently joined them....for no viable reason.
Seriously, they were already in the apple app store, they only do apple ports, and the only machines that could run them all had the store already installed. Why Feral would join EA in anything is beyond me, but it concerns me.
Personally, I wouldn't buy anything through Origin because it is an EA product, and EA has done terrible things to the industry. If you can get the same game elsewhere, do so.