The logic boards in the Mac Pros are of proprietary Apple designs, and built at Foxconn.
A 2009 (4,1) board should support any CPU with D0 stepping (3500/5500 series), but would require a flash to 5,1 firmware to support processors with B1 stepping (3600/5600 series).
Good info, thanks for that. I'm getting closer to committing to a CPU upgrade now