While in the past this was always true - the fact that you need to install Snow Leopard first and then install Lion over it when you do a clean install. Opens the door slightly for a chc at the New Mac Pro to be able to run Snow Leopard.
You don't actually. You need to have the latest Snow Leopard installed if you buy Lion from the App Store the regular way, but once you have the package, you can just go do clean installs on any box you want.
Edu and multiple seat licenses for larger corps, will work the same as before (physical media, clean installs, or that's how it looks
right now).
Unless Apple radically changes their methodology, the oldest OS version that will run on your boxes, is the one that comes with it when you buy it (as you pointed out).
For new hardware that's usually an unreleased version of whatever the current point release happens to be, with new drivers for the hardware in question; which will make the "standard" build, the next update.
In other words, if for the sake of argument you buy a new Mac, while everybody is on 10.6.5, you will have an unreleased version of OS/X (including your install/backup media), which contains 10.6.5 with a different build #. By the time 10.6.6 rolls around, all those drivers will be part of the mainstream build.
If your new computer comes with Lion, it will probably not be running Snow Leopard unless you install it on a VM. If Apple is indeed holding up release of new hardware like the new MacBook Air in particular, waiting on Lion, then you can assume that they really want everybody on Lion as fast as possible and may go out of their way to make any kext transplants very difficult or impossible, to backport to older versions of OS/X.
FCP 7 will continue to work on Lion, according to Apple's official press release circa yesterday. That one was possibly not planned on, but became officially supported due to the uproar amongst pros who were using Apple's, what is it again, oh right, "pro" software and can't just drop everything and wait on FCP X to catch up on all the features that got left out.
Being unable to run Rosetta on the other hand, isn't going to make anybody storm the Apple mothership with pitchforks. It's more like, oh well, it's been a realllllllly long time now, if you still haven't upgraded or have some ancient legacy software, try to get over it, the world has moved on.