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It's true that prior to the 2012 Mac Pro, Apple was not only price competitive at the high end (the fat), but I found them them to be less expensive than their greedy competitors who were and are on fatfull diets.
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Nope. Neither you nor I know what the exact computational potential of the low end box will be, but we can use history as a guide (and the base has been watered down). This is the area where Apple has not been price competitive traditionally, .... I acknowledging that Apple sold more units at the low end at higher prices than its competitors.
They sold more at the low end with a box not optimized to sell at the lower end of that spectrum. This one is. It walks away from the dual processor market so it is more much more a single CPU focused offering. That doesn't necessarily be loss in computational horsepower, just fewer x86 cores.
I don't think Apple has any keen interest in selling $5-12K boxes to stagnant group of customers. The other system vendors like the high priced boxes because they can goose the margins. Typically those increased margins are used to offset "loss leader" boxes they have elsewhere in their line up. Apple has no loss leader boxes. That is one reason why their prices were more reasonable as go to the top end of the scale.
This new box should move the top end of the scale to a new place. Again Apple has little to no need to goose standard configuration prices increasingly higher because not in a "rob Peter to pay Paul" exercise.
As far as history goes Apple has historically put a technology based performance gap between products and their pricing. I don't think Apple is likely to shrink the GPU in the entry Mac Pro so much that it is below the where the higher iMac BTO options are. So a W7000 equiv is probably as low as they go. But yeah that is a educated guess.
One of the problems Apple had with the single CPU package box is that it needed to be higher than iMac and lower than the dual. If they toss the upper end they will have more freedom to put value-add into the entry single that won't wipe out dual sales. Dual sales are off the table.
If left with just needs to be higher than iMac there is little reason to no stuff move value ( e.g., multiple TFLOPs of computational horsepower) so that the $2,499 is buying alot more. Paying $2,499 for empty space where can later put something is pretty expensive empty space. It think it is going to be much easier for Apple to sell an actual thing (here is the 2nd GPU this is what it can do ) than "empty space".
Isn't the true reason why a true upgrade from Mac Pro 2010 has taken so long is that Apple wasn't satisfied with the tiny share of the overall market that it was eating?
Time will tell if this was a "cut loose the dead weight, so can maneuver better" move to better serve the single CPU but moderately high core count market or " the discrete GPU card market spawn a resurgence in boxes with slots".
Personally, I would not bet again some form of GPGPU being folded into CPU packages in the Xeon E5 line up in the future. It will become commonplace for there to be at least one GPU that is embedded into the system. Same thing in entry/mid-level server market ( 1 CPU package) .
The "attack of the killer micros" killed off practically all of the specialized supercomputer CPUs from 2 decades ago. The same forces will kill off the the dual / quad micros over time using same technology forces. For example the Xeon E7 line up is starting to be endangered by the E5 4600 line up. E7 will shrink to smaller number of boxes that are death sprial kind of pricing ( higher price ,, fewer customers, higher price ,,, ... )
Over time computers have generally gotten smaller. Not only Apple's ... all system vendors. You'd be hard pressed to name a single successful system vendor who did better long term by "going bigger" over time rather than smaller. In fact, many of the industry inflection points have "big" vendors dragging their feet on going small because catering to the legacy , sunk cost market that was stuck on the larger form factor.