My guess is:
Q1-Q2 2016
Intel Xeon E5 v4 (Broadwell EP)
DDR4-2400 RAM (16GB min)
New PCI-e 3.0 SSD (256GB min, but cheaper 512GB-1TB options)
Double internal slot for SSD (1 left empty in the 256GB version)
6 Thunderbolt 3/USB 3.1 ports
Unless dumped the other GPU, the E5 v4 runs out of PCIe v3 lanes after get to two x4 PCIe v3 TB3 controllers. So no v3 SSD. No 3rd controller to get to 6 TB sockets either. Can't crank up both SSD and TB. It is going to be one or the other.
The C612 chipset supports 6 USB 3.0 sockets so maybe 6 of those, but probably just 4 of the new TB flavor.
Second SSD slot is doubtful since same x8 PCIe v2 limit the current Mac Pro has. Apple could put in a switch so the two x4 PCIe v2 SSDs can share the same x4 PCIe v2 allocation. That buys capacity by trading off speed. If try ti read or write at some time to both then get half the throughput. I suspect most folks are going to want top end speed from both. If they go to NVMe SSD that is even worse as the individual SSDs will be even closer to maxing out the x4 link all by themselves; let alone 2.
Apple could back away from the "More than two" TB socket path. One TB v3 controller. Some other Type C sockets that are just just DP+USB 3.0. Still have max 6 mainstream displays but it turned out folks didn't often attach 4+ TB devices to the systems. That would free up a x4 PCIe v3 link on CPU's lanes for ultra fast SSD connection. The other 2nd slot would be a just a 4x PCIe v2 SSD. Apple would have to get over their OCD symmetry syndrome though.
My secret hope is to see some "Jobs style" anticipated purchase of Skylake EP processors, like they did with the first Xeon (first Intel Mac Pro). But I think we need to see whether Intel wants to skip the Broadwell EP generation.
There was likely no "Jobs style" anticipated purchase ... in 2009.
Most reasonable large system makers that are building Xeon server class system gets get some "early" (as in before initial announcement) in some reasonable volume. That is so they can roll out "test"/"qualification" servers under NDA to their larger customers ( who typically have long purchase request timelines) and to ramp production on.
Apple merely jumped out the gate early with that initial drop. Relatively few beta units sent out and not a top 5 volume players so probably had enough with lower production targets to ramp to.
Apple didn't have special access they just ran out of the gate early whereas the other players we waiting to sync up with Intel's official announcement. It was more Apple pissing on Intel's launch marketing event than special access thing. It wasn't a norm. It still isn't one.