thanks for that depressing realisation that my 2009 iMac has SIGNIFICANTLY been outdated
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Take heart. After installing the OS X update on the the 2010 iMac Core i7, I cloned its HDD to an external HDD. Then I booted the 2009 iMac Core i7 with that drive.
Portal at 2560x1400 "High" with 4X MSAA
2009 iMac Core i7 Before = 19 fps
2009 iMac Core i7 with Cloned Drive = 42 fps
2010 iMac Core i7 = 59 fps
ETQW 2560x1400 "High" with 4X MSAA
2009 iMac Core i7 Before = 29 fps
2009 iMac Core i7 with Cloned Drive = 44 fps
2010 iMac Core i7 = 48 fps
And X-Plane 2560x1400 "High" with 2X MSAA
2009 iMac Core i7 Before = 80 fps
2009 iMac Core i7 with Cloned Drive = 114 fps
2010 iMac Core i7 = 128 fps
WoW went from 42 fps to 49 fps on the 2009 (vs 52 fps on 2010)
Team Fortress went from 29 fps to 44 fps on the 2009 (vs 44 fps on 2010)
These numbers confirm to me that the newer drivers account for part of the gains. The 2010 iMac Core i7 is still faster, though not as dramatically.